Have you tried opening a Case with Now Support (formerly known as HI)?  They should be able to take a look at your instance and resolve this for you. 

There are two things I would check.

1. Does the user doing the submission in Dev have an email address defined on their User ID?  I don't think it applies to the error message above, but that is a requirement that can cause submissions to fail. 
   
   
2. In your Controller (should be Prod) open the "All" Nav menu and go to "App Engine" >> "Pipelines and Deployments" >> "Pipeline Types". 
1. There should be a single record there named "Application Deployment". 
2. Open that record. 
3. Scroll down to the related list at the bottom named "Pipeline Instances".
4. Sort this list by updated. 
5. Personalize the list layout and add the column "Pipeline Validation Details".
6. This list shows the result every time someone attempted to "Submit" an application and the "Pipeline Validation Details" column can contain critical information for determining what may be causing the issue. 
7. Here is an example screenshot of one of the records showing where the User submitting in Dev does not have an email address defined for their User ID.